The Pfam:tRNA-synt_1c_C domain within your query sequence starts at position 396 and ends at position 583, and its E-value is 4.3e-48.

APRAMAVLEPLQVVITNFPAPKPLDIRVPNFPADETKGFHQVPFASTVFIERSDFKEESEPGYKRLASGQPVGLRHTGYVIELQNIVRGSSGCVERLEVTCRRADAGEKPKAFIHWVSQPLVCEIRLYECLFQHKNPEDPVEVPGGFLSDLNPASLQVVEGALVDCSVALAKPFDKFQFERLGYFSVD
Pfam:tRNA-synt_1c_C

Pfam:tRNA-synt_1c_C
Pfam domain

Pfam ACC: