The domain within your query sequence starts at position 19 and ends at position 371; the E-value for the SPESP1 domain shown below is 5.8e-155.

YPTITVLPDEEQNLNHYVHILQNLIMSVPTKEQDLGKKLSSSRTVDSAEPRSLASKVLLT
PGLVSAQDVTPESDVLIRPVDETTNSRTRGFTLRRRRTQSTAFWSIRPNNISVVLRTEEP
FIEKEPEPELESSRLPTEPEPELEPEPEPVAESRQMSEPEEELVTSTTPNKELTGTSRIS
SMATQPANTQATRITVTVKTTSTMDVSTDSEDVPQLSGQSEIPSAEDLPGRHSLNTRHED
ILKKISNINAEIQQGLLGGNNSPEFKEFIKASREHLKRSLALAAAAEHKLEQMYGSNVFP
EGRTSDPDNDMEMIINMLYNSRSKLSDYFNIKRVPSELREKASVVNAELRKIL

SPESP1

SPESP1
PFAM accession number:PF15754
Interpro abstract (IPR026743):

Equatorial segment protein (ESP) been localised to the equatorial segment of the acrosome in sperm. It defines a discrete acrosomal subcompartment that persists throughout acrosomal biogenesis [ (PUBMED:12773409) ] and is required for fully fertile sperm [ (PUBMED:20375058) ].

GO process:acrosome reaction (GO:0007340), fusion of sperm to egg plasma membrane involved in single fertilization (GO:0007342)
GO component:acrosomal vesicle (GO:0001669)

This is a PFAM domain. For full annotation and more information, please see the PFAM entry SPESP1