The domain within your query sequence starts at position 1387 and ends at position 1440; the E-value for the RPT_1 domain shown below is 6.48e-5.

PEQGGESVESTKCQVVSKSTENCEDKLQGKVTEANGKKLGQHPPKPEERAVNRC

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.