The domain within your query sequence starts at position 1482 and ends at position 1554; the E-value for the RPT_1 domain shown below is 2.88e-9.

NLVDPGSSPSQVLSAATIVAKHTSALCNACRIASSKTANPVAKRHFVQSAKEVANSTANL
VKTIKALDGDFSE

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.