The domain within your query sequence starts at position 286 and ends at position 472; the E-value for the RPT_1 domain shown below is 4.17e-7.

SPSSSRSPCHNLSVSLTSDSNVHLNSNAHSIQSSGAQVPYVPSATGFSNVRVPLMPSKTV
SCSSIAPHMSLIKVPSSIQDLHLPTQEAFSSTEDPHNQAIASKNLQTTKMPPTTLTSKAQ
LLKMPSSATSSSSVQVSHTLATVSRSTSVKQILQTTRSSSIQILNSATVKASKNVQASQV
SLPTEPN

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.