The domain within your query sequence starts at position 326 and ends at position 414; the E-value for the RPT_1 domain shown below is 1.39e-5.

KSLQSLYREHCEAILDVVVNLQFSLIEKLWQTFWRYSPSTPADGTTITESSNLSEIESRL
PKAKLITLCKHESILKWMCNCDHGMYQAL

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.