The domain within your query sequence starts at position 464 and ends at position 552; the E-value for the RPT_1 domain shown below is 1.82e-5.

KSLEGWLSNAMNNIPQRMIQTKVAAVSAFAQTLRRYTSLNHLAQAARAVLQNTSQINQML
SDLNRVDFANVQEQASWVCQCDDNMVQRL

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.