The domain within your query sequence starts at position 467 and ends at position 556; the E-value for the RPT_1 domain shown below is 1.35e-6.

SEQLTPSSATFPDPLTSSLQGQLTESSARSFEDQLTPCTSSFPDQLLPSTATFPEPLGSP
AHEQLTPPSTAFQAHLNSPSQTFPEQLSPN

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.