The domain within your query sequence starts at position 473 and ends at position 570; the E-value for the RPT_1 domain shown below is 7.22e-14.

RQPAGEANPLIGSLLQERQDVIARIAQHLEHIDPAAHIPRPAFSKHDSNSIPSKVFRSSF
DDKTLLKKGRENVSVSVSHTEVSLLGDRGDGESLKPSK

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.