The domain within your query sequence starts at position 503 and ends at position 779; the E-value for the RPT_1 domain shown below is 3.34e-9.

MKVPEMKLPKVPEMAVPDVHLPDVQLPKVPEMKLPKVPEMAVPDVHLPDVQLPKVPEMKL
PEMKLPKVPEMAVPDVRLPEVQLPKVSEVKLPKMPEMAVPDVHLPELQLPKMSEVKLPKM
PEMAVPDVRLPEVQLPKVSEMKLPKMPEMTMPDIRLPEVQLPKVPDIKLPEMKLPEIKLP
KVPDMAVPDVPLPELQLPKVSDIRLPEMQVSQVPEVQLPKMPEMKLSKVPEVQRKSAGAE
QAKGTEFSFKLPKMTMPKLGKVGKPGEASIEVPDKLM

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.