The domain within your query sequence starts at position 537 and ends at position 652; the E-value for the RPT_1 domain shown below is 3.02e-34.

PKKIMAPPERDMSPPEGDVAPPKKIMAPPERDMSPSEGDVAPPKKIMAPPERDMSPPEGD
VAPPKKIMAPPERDLSPSEGDVAPPKRIMAPPERDMSPFKGDMAPPKGIMEPPNRD

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.