The domain within your query sequence starts at position 56 and ends at position 459; the E-value for the RPT_1 domain shown below is 4.41e-9.

TSASDLEGTSTQLMTSPGFDSLSTPLMGAPHSGTLSPPLMSASDSGTLSPLLMPASDSGT
LSPLLMPVSDSGTLSPLLMPASDSGTLSPLLSTSDYGLMSPGMMSIPDFGTMSSLMAAPD
SAEISPLAMPIQSSGVISAPIMSTSSSEASLMLGSDPGEISPLLIPDMNPGVTSTPPMTA
PGSEAMSPLQITDEDTEAMSKVLMTALASGEISSLLMSGTDSEAISSLIMSALASGETPA
QPTNPPESEGIPTVLMSGSDSAVMSSLPMPVSGSGAMPTPLLSIPDAGEIATLPKPVPDV
EAMSPLLMTALTSTVMPSQLISASSSGVMSPDTTQNINSEVMSAPPIRVSSTGLMSTLPV
RASDTAATPIQLMRVPASGNMSTSQKTVPVSGSMSTPLMAVTSP

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.