The domain within your query sequence starts at position 315 and ends at position 360; the E-value for the RPT_2 domain shown below is 1.18e-9.

GPQGPLGYPGPRGVKGVDGIRGLKGHKGEKGEDGFPGFKGDIGVKG

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.