The domain within your query sequence starts at position 485 and ends at position 555; the E-value for the RPT_2 domain shown below is 1.1e-9.

RSRSKERDHETTKEKEKPLDPKGKDQERSRSKENSKQVESKSNEHDHSKSKEKDRRAQSR
SRERDLTKSKH

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.