The domain within your query sequence starts at position 593 and ends at position 707; the E-value for the RPT_2 domain shown below is 6.03e-6.

QVRGEYTVDISTLESDQREGGDADAANQVTPEAEEQARSSEPDMETCIVPSQDEEKTEEE
NGMYALAQGKRNLRGANGTDGAQTQKLGEENQGQLGNESHKMIHGPKRKNQKQVR

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.