The domain within your query sequence starts at position 1412 and ends at position 1470; the E-value for the RPT_3 domain shown below is 2.21e-5.

EKFSCLQSEHKILHDQHCEVSSKMSALRSYVDTLKAENSALSMSLRTLQGDLVKEGEPA

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.