The domain within your query sequence starts at position 621 and ends at position 885; the E-value for the RPT_4 domain shown below is 2.09e-6.

LETVAHELQRESQEINLPNQKNTSLGDMNCESHGPEVGFHADAQLEADQSPVNPEEVDLD
LENQSVHSGISNLSFDSNASYQSANDQPQGAWGEVNLDELNVDMEVKSNGCSSSELTFDS
DSPLLSVTERSLLDFEGLNEDDFNLEDENCVSSSSDITFDSDIPDDSVADQPQVAVYEEE
PVGLENKSNESCVSGITFDSDIPLHSGNDHPEVAVKEVIIKEDENVHLEGKNDNPSGSEI
CLDSNVPLHSVTNSDVAVKKINPPK

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.