The domain within your query sequence starts at position 690 and ends at position 739; the E-value for the RPT_4 domain shown below is 4.11e-8.

GDTGQKGLPGPPGPPGYGSQGIKGEQGPQGFPGSKGTVGLGLPGQKGEHG

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.