The domain within your query sequence starts at position 407 and ends at position 565; the E-value for the RPT_6 domain shown below is 7.68e-5.

VDQEDASYESRGSDMSFDCGSSCQSLSALSELTAREINVSEETHAYSQPRNETPTVSGAT
SDNGSSSRQVITQNIQLISLVDESYESSGSEVNFDGDDSLPSTSHRPPQPVEVVVPLRLV
DKSYGSGSSEPCEDSGSSADETPAASRQQNPVRNTHANL

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.