The domain within your query sequence starts at position 373 and ends at position 534; the E-value for the RPT_1 domain shown below is 5.23e-7.

PTSPESWKSGPPELRKTALPLSPEHWKAVPPVSPELRRPGPPLSPEIRSPAGSPELKKPS
SSPDLWKVSPDQRKTSPASLDFPEPQKSSCGSPPDLWKSSFIMESQKPNVFSETRKHTAS
GSSESPKVASDIWKPVLSIDAEPRKSTLFPEPTKAVLPASPE

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.