The domain within your query sequence starts at position 390 and ends at position 540; the E-value for the RPT_1 domain shown below is 9.24e-13.

DNPRTPCTQRDNPRASSPNRTTQRDNPRTPCTQRDNPRASSPNRTTQRDNPRTPCAQRDN
PRAASPNRSTQRDSPRTPCAQRDNPRASSPNRTAQRDNPRTPCAQRDNPRTSCTSQNTPR
TPSTQADKTTASCSKWEHLRSACTQRDNPRT

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.