The domain within your query sequence starts at position 457 and ends at position 643; the E-value for the RPT_1 domain shown below is 2.14e-7.

GKGDPQPWKKASLPASEKTDPLASSKAGSASQGKAETVSPGEVDAMTLGKTVPTSSGKTA
LVSPGKVDLMTSERAEGIPELQASEKGNPVNSTRVDTGATGSTEPKSGVKVITQIPGATS
PGKVETPSLQKEQPQLSEKTDPSRKVDPPTTVEPVSLGKADSASPSPRKAESQTSAKTVP
QAPDKAT

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.