The domain within your query sequence starts at position 471 and ends at position 675; the E-value for the RPT_1 domain shown below is 3.72e-6.

VASQASQGTGSPIPKIHGSSFLTSTVKQEESLFASMPPLCPIGSHPKVQSPKAVTGGLGA
FTKVIIKQEPGEAPHVSTTGAASQSAFPQYVTVKGGHMIAVSPQKQVISAGEGTTQSPKI
APSKVVGVPVGSALPSTVKQAVAISSGQILVAKASSSVTKAVGPKQVVTQGVAKAIVSGG
GGTIVAQPVQTLTKTQVTAAGPQKS

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.