The domain within your query sequence starts at position 472 and ends at position 573; the E-value for the RPT_1 domain shown below is 2.14e-5.

NCGFSQGNGDTVNSALSAKQNGSSSAVQKEGNGGNAWDSGPPAGPGILAWGRGSGTNGIG
NIHSGAWGHPSRSSSNGVNGEWGKPPNQHSSSDISGKGSTGW

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.