The domain within your query sequence starts at position 485 and ends at position 695; the E-value for the RPT_1 domain shown below is 4.85e-5.

LLTPDATTQNNKPQVGEGTKERLENSDSSTTETTAVQVLSEPMEKADITNHVFATSAVGV
NTPAEASPALSSEEIPTEKPGMETQERGCEGGTTSDQSSPVLPAAAIENKVLGGQEPDTS
IAGFCKTASPLDLTMPGPSSDGMPEQNSESHARPAQSLSGQALLCSTAEAGTPSAEATHQ
PSTVTSSGRLEECGSGKASLPESTMVQPSTQ

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.