The domain within your query sequence starts at position 507 and ends at position 750; the E-value for the RPT_1 domain shown below is 2.93e-7.

EFEQPVAMTTVEHPGHPEVTTATGLLGQPEAAMVLELPGQPVATTALELSGQPSVTGVPE
LSGLPSATRALELSGQSVATGALELPGQLMATGALEFSGQSGAAGALELLGQPLATGVLE
LPGQPGAPELPGQPVATVALEISVQSVVTTSELSTMTVSQSLEVPSTTALESYNTVAQEL
PTTLVGETSVTVGVDPLMAQESHMLASNTMETHMLASNTMDSQMLASNTMDSQMLASNTM
DSQM

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.