The domain within your query sequence starts at position 569 and ends at position 638; the E-value for the RPT_1 domain shown below is 9.92e-6.

SSSQKASSEGLSKGKKTANGLAADANFSNSKAPILSKYPYSSRSQENILQGFSLPNKEGV
DQPVAVVMED

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.