The domain within your query sequence starts at position 218 and ends at position 373; the E-value for the RPT_2 domain shown below is 2.94e-114.

GKKGEGAQNQAKKGEGAQNQAKKGEGAQNQGKKGEGAQNQAKKGEGGQNQAKKGEGAQNQ
GKKGEGAQNQGKKGEGAQNQAKKGEGAQNQAKKGEGAQNQGKKGEGAQNQSKKGEGAQNQ
AKKGEGGQNQAKKGEGAQNQAKKGEGAQNQAKKGEG

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.