The domain within your query sequence starts at position 4549 and ends at position 4697; the E-value for the RPT_2 domain shown below is 6.03e-7.

FLLSGCCQALEISLDQEHLPFGPVVYQTQATRRILIMNTGDVGARFKWDVKKLKPHFSIS
PEEGYIISGTEVALEVTYHPTEIGKESLYKNILCFIQGGNPLCLTLSGTCVGPPVVKEVV
NFNCQVRSRHTQTILLSNRSNQTWNLHPI

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.