The domain within your query sequence starts at position 479 and ends at position 581; the E-value for the RPT_2 domain shown below is 3.9e-5.

GSGVERMGAGMGFGLERMAAPIDRVGQTIERMGSGVERMGPAIERMGLSMDRMVPTGMGA
SLERMGPVMDRMATGLERMGANNLERMGLERMGANSLERMGLE

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.