The domain within your query sequence starts at position 514 and ends at position 568; the E-value for the RPT_2 domain shown below is 5.51e-5.

VPKYRTPKERVSYHKASKDVSPEDTGTKDEAPEVKVLKKVVYKDRPQKDKAPEPR

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.