The domain within your query sequence starts at position 575 and ends at position 683; the E-value for the RPT_2 domain shown below is 6e-5.

NEELNCQDPDVGDMEEEERLQVTDKKQASGSPEQKIRAGSREPVEDPQSGSSGKQNKVEE
DGPTEGPTDILDQNSPQCEDREISPVGEKGPQCDTSQIGSEEGHVTSQH

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.