The domain within your query sequence starts at position 316 and ends at position 419; the E-value for the RPT_3 domain shown below is 5.13e-5.

EGAAVVKVPTLDVAAPSMGVDLALPGAEVEAQGEVPEVALKMPRLSFPRFGIRGKEATEA
KVVKGSPEAKAKGPRLRMPTFGLSLLEPRPSGPEAVAESKLKLP

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.