The domain within your query sequence starts at position 1335 and ends at position 1386; the E-value for the RPT_6 domain shown below is 4.17e-5.

ESSEGEEHSVVPQRNSESCHCHCHDQAGHQQRESVHGQRGRPEGPSQDSSRH

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.