The Pfam:tRNA-synt_1e domain within your query sequence starts at position 319 and ends at position 387, and its E-value is 3.4e-6.

GAGLRPPHRIYVHSHWTVSGQKMSKSLGNVVDPRTCLDRYTVDGFRYFLLRQGVPNWDCDYYDEKVVKL
Pfam:tRNA-synt_1e

Pfam:tRNA-synt_1e
Pfam domain

Pfam ACC: