The domain within your query sequence starts at position 311 and ends at position 408; the E-value for the RPT_1 domain shown below is 1.8e-29.

LRDQEERAQAEAVGSPGTSAKDKAEAKVTLVDSSCKYQAESKVTLVDSSCKDQAESKVTL
VDPSCKDNAKSKVTLGSSRKYKAKSKVPLVDSSCKDQA

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.