The domain within your query sequence starts at position 323 and ends at position 439; the E-value for the RPT_1 domain shown below is 3.7e-11.

PGPRGVKGVDGIRGLKGHKGEKGEDGFPGFKGDIGVKGDRGEVGVPGSRGEDGPEGPKGR
TGPTGDPGPTGLMGEKGKLGVPGLPGYPGRQGPKGSLGFPGFPGASGEKGARGLSGK

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.