The domain within your query sequence starts at position 402 and ends at position 510; the E-value for the RPT_1 domain shown below is 8.05e-5.

YDGQIAVFGTDLQEKLSDQHYLLVGAGAIGCEMLKVFALVGLGVRANGGVTVADMDYIER
SNLSRQFLFRPKDVRRPKAEVAAAAAHRLNPDLRATPYTCPLDPTTEDI

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.