The domain within your query sequence starts at position 413 and ends at position 597; the E-value for the RPT_1 domain shown below is 2.35e-83.

HVDLKGPKVDVKGAKGEVSAPDLEVSLPGVEVDIQAPGAKLEGELSLADKDVAAKDSKFK
MPKFKMPSFGVSAPGKPSLEASLEVGAPKVEADVAPPQVQGDLKTPDLSVQLRSADLELK
AGQVGVKLPEGQLPEAELPAAQAAAAAGAGLKGHLPKVQMPSLKMPKVDLKGPHVDLKGP
KVDVK

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.