The domain within your query sequence starts at position 438 and ends at position 538; the E-value for the RPT_1 domain shown below is 8.15e-5.

PDLPSRENAKDELVSLLAPLEEKKPQMKEFPVTEAAKFQTDVKLEREGVLEFEEKGLEIC
QDSNLEAAESTESYSILRFAKPDVSDPEPTARKDEYKLDTK

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.