The domain within your query sequence starts at position 439 and ends at position 550; the E-value for the RPT_1 domain shown below is 4.96e-7.

ERPLVTDMTPSLETEMTLGKETAPPTETNLGMAKDMSPLPESEVTLGKDVVILPETKVAE
FNNVTPLSEEEVTSVKDMSPSAETEAPLAKNADLHSGTELIVDNSMAPASDL

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.