The domain within your query sequence starts at position 542 and ends at position 659; the E-value for the RPT_1 domain shown below is 4.88e-5.

EKLLKKQKVPAALLEVTVADTEPTQTQTLLRSLGELVTGIYVKYWIYVCAGMFIVVSFAG
RLVVYKIVYMFLFLLCLTLFQVYYTLWRKLLRVFWWLVVAYTMLVLIAVYTFQFQDFP

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.