The domain within your query sequence starts at position 575 and ends at position 644; the E-value for the RPT_1 domain shown below is 2.5e-5.

PGEQAPRELAEDEPSTDRSAPVNGEATSQKGENMEDRAQEDGQDSEDGPRGGSSEELHDP
RDNSDPAKPG

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.