The domain within your query sequence starts at position 399 and ends at position 595; the E-value for the RPT_2 domain shown below is 4.46e-7.

QPGMKYSESGPWSPRHLDMRQPSPSQLATRQFDKWQQNPSMPGVRQLYTWQPTSSFSDTR
QLEKFHPCPINLNMEQFWEAVSRQPGTTQLGTNQLDTNQPDGTQSSQGGKTQSDKLEPSP
RKPEMKGSQPDTSQSDSDHLDISQPGPSQLEPGESSMSDLNESQQRITQSPMGKKDSCSF
FIRPAEPEDCPDILRLI

This is a significant internal repeat in the query protein sequence, as detected by the Prospero program.